Alcobaça Formation

The Alcobaça Formation, previously known as the Guimarota Formation, is a geological formation in Portugal. It dates back to the Oxfordian stage of the Late Jurassic.[1] It is an important source of information on the diversity of Late Jurassic mammals.[2][3] Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.

Alcobaça Formation
Stratigraphic range: Oxfordian-Tithonian
~160–145 Ma
TypeGeological formation
Sub-unitsCamadas de Guimarota
UnderliesLourinhã Formation
OverliesMontejunto Formation
Lithology
PrimaryMarl
OtherMudstone, coal, siltstone, limestone, sandstone
Location
Coordinates39.6°N 9.0°W / 39.6; -9.0
Approximate paleocoordinates33.7°N 1.6°E / 33.7; 1.6
RegionEstremadura, Leiria, Lisboa
Country Portugal
ExtentLusitanian Basin
Type section
Named forAlcobaça, Portugal
Named byChoffat
Year defined1885
Alcobaça Formation (Portugal)

Vertebrate paleofauna

Dinosaur eggs are geographically located in Lisbon District, Portugal.[4] Dinosaur tracks are geographically located in Leiria District, Portugal.[5]

Amphibians
Amphibians of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Celtedens Indeterminate Porto Pinheiro Albanerpetontid
Choristoderes of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Cteniogenys Indeterminate Guimarota

Ornithischians

Indeterminate euornithopod remains located in Lisbon District.[4] Indeterminate stegosaurid remains present in Lisbon District.[4]

Ornithischians of the Alcobaça Formation
GenusSpeciesDistrictStratigraphic positionAbundanceNotesImages
Dacentrurus[4] D. armatus[4] Lisboa[4]
Ornithopoda[4] Indeterminate[4] Lisboa[4]
Trimucrodon[4] T. cuneatus[4] Lisboa[4] Later determined to be indeterminate ornithischian remains.[4]

Saurischians

Indeterminate sauropod remains located in Leiria and Lisboa.[6]

Saurischians of the Alcobaça Formation
GenusSpeciesDistrictStratigraphic positionAbundanceNotesImages
Allosaurus[5] Indeterminate[5] Leiria[5]
Aviatyrannis A. jurassica Guimarota
Lusotitan[4] L. atalaiensis[4] Lisboa[4]
Ceratosaurus[4] Indeterminate[4] Lisboa[4]
Dinheirosaurus[4] D. lourinhanensis[4] Lisboa[4] "[Two] cervicals, [nine] dorsals, ribs, limb elements."[7]
Lourinhasaurus[4] L. alenquerensis[4] Lisboa[4]
Megalosaurus[4] M. insignis[4] Lisboa[4] Later determined to be indeterminate theropod remains.[4]

Turtles
Turtles of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Peltochelys P. duchastelli Guimarota
Platychelys Indeterminate Guimarota
Pleurosternidae Indeterminate Guimarota

Lepidosauromorphs
Lepidosauromorphs of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Becklesius B. hoffstetteri Guimarota
Dorsetisaurus D. purbeckensis Guimarota
Marmoretta Indeterminate Guimarota Basal Lepidosauromorph
Portugalophis P. lignites Guimarota Stem-snake
Saurillodon S. proraformis Guimarota
Saurillus S. henkeli, cf. obtusus Guimarota

Crocodyliformes
Crocodyliformes of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Goniopholis G. baryglyphaeus Guimarota
Knoetschkesuchus K. guimarotae Guimarota
Lisboasaurus L. mitracostatus Guimarota
Machimosaurus M. hugii Guimarota

Mammals
Mammals of the Alcobaça Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Bathmochoffatia B. hapax Guimarota
Bernardodon B. sp, B. atlanticus Porto Pinheiro
Drescheratherium D. acutum Guimarota
Dryolestes D. leirensis Guimarota
Ecprepaulax E. anomala Porto Pinheiro
Guimarotodon G. leiriensis Guimarota
Guimarotodus G. inflatus Guimarota Dryolestidae
Haldanodon H. exspectatus Guimarota
Henkelodon H. naias, H. sp Guimarota
Henkelotherium H. guimarotae Guimarota
Iberodon I. quadrituberculatus Porto Pinheiro
Kielanodon K. hopsoni Guimarota
Krebsotherium K. lusitanicum Guimarota
Kuehneodon K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp Guimarota
Laolestes L. andresi Porto Pinheiro
Meketibolodon M. robustus Guimarota
Meketichoffatia M. krausei, M. sp Guimarota
Nanolestes N. krusati Porto Pinheiro
N. drescherae Guimarota
Paulchoffatia P. delgadoi Guimarota
Pinheirodon P. sp, P. vastus, P. pygmaeus Porto Pinheiro
Plesiochoffatia P. thoas, P. peperethos Guimarota
Portopinheirodon P. asymmetricus Porto Pinheiro A dryolestid
Priacodon Indeterminate Porto Pinheiro
Proalbionbaatar P. plagiocyrtus Guimarota
Pseudobolodon P. dryas, P krebsi, P. oreas Guimarota
Renatodon R. amalthea Guimarota A paulchoffatiid
? Tinodon Indeterminate Porto Pinheiro
Xenachoffatia X. oinopion Guimarota
Paulchoffatiidae Indeterminate Guimarota 4 distinct genera with 7 species
Pinheirodontidae Indeterminate Porto Pinheiro
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
 Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

References
  1. Weishampel et al., 2004, pp.545–549
  2. "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
  3. "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
  4. "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
  5. "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
  6. "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
  7. "Table 13.1," in Weishampel, et al. (2004). Page 265.

Bibliography
  • Weishampel, David B.; Peter Dodson, and Halszka (eds.) Osmólska. 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21. ISBN 0-520-24209-2
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